The oligocene berwick quarry flora — rainforest in а drying environment

The Late Oligocene to possibly earliest Early Miocene Berwick Quarry macrofossil flora was first described very early in this century by Henry Deane, but has since been largely ignored. Recent work at the quarry has led to major new collections and а reinvestigation of the flora. Seventeen taxa of macrofossils have been recovered, including Agathis, Dacrycarpus, four species of Lauraceae, Gymnostoma, Nothofagus, Eucalyptus, an indeterminate Myrtaceae and Proteaceae, three possible Cunoniaceae, and six unidentified taxa. Fagus maideni Deane is formally transferred to Nothofagus Blume. Nothofagus johnstoni Hili, Fagus luemanni Deane, and Atherosperma berwickense Deane, are considered to be junior synonyms of the species N. maideni (Deane) comb. nov. Only one of the seven fossil species of Eucalyptus established by Deane can be recognised in our material:E kitsoni Deane. His Lomatia species are probably Cunoniaceae. The Berwick Quarry flora contains species which are consistent with rainforest in the region, however, the flora is also notable for the presence of leaves of Eucalyptus, leaving little doubt that the flora at Berwick Quarry represents а mixture of rainforest and open forest taxa. The vegetation was probably а mosaic of open and closed forest, representing some of the earliest evidence for seasonality in Australia during the development of the modem flora. Palynological samples are dominated by Nothofagus subgenus Brassospora and Fuscospora.